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Growth and survival can be split into specific phases depending on the yearly events of the life cycle in the two time steps (summer and winter). The computational order of life cycle events and processes, together with their length (in days) when relevant, are presented in the figure above.
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## Process overview
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In the river phase, individuals grew in weight according to individual and stage-dependent growth capacity and influenced by water temperature, population density and river flow. Growth increments in weight were then allocated to fat reserves (Fat) or somatic growth through an increase in body length depending on a variable individual propensity to accumulate fat. Survival in the river was phase dependent, with higher mortality for maturing individuals and during winter. The triggering of sea migration was size dependent 6 months before the run. The smoltification process allows an individual that was in the river (‘parr’) to become physiologically ready to run into the sea (as ‘smolt’). The probability of smolting for an individual followed a reaction norm based on its body length.
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